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Identification of Canada and
Cackling Goose |
October 7, 2004 |
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One of the most significant changes for birders in the 45th supplement of the AOU checklist (Banks et al 2004) was the split of Canada Goose into two species. While widely anticipated - the highly variable Canada Goose has often been considered more than one species in the past - the split has thrust a complex and poorly understood field identification problem into the forefront. I have tried to pull together some of the basic information about subspecies and variation to begin the approach to identification of these species. I must stress that this is a draft, and should not be considered anything more than my personal notes on the problem of Goose identification. It stresses the problems in eastern North America, and includes lots of unanswered questions. I hope to add more detail in the future. Please direct any comments or discussion to me or to the ID-Frontiers listserv. What is the
"Cackling Goose"? When reading published accounts of subspecies, it is important to bear in mind that in the confusing history of the taxonomy of Canada Geese, the subspecies have been grouped and named in various ways. Names such as leucopareia (which is now restricted to the Aleutian population) have been applied to much broader groups in the past. The names attached to individual specimens in past decades might not be the same names that would be used in the current subspecific arrangement… beware. The subspecies
defined below should be taken cautiously as merely one interpretation of
the information at hand (which is patchy and incomplete, despite decades
of study). |
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| Subspecies - See map figure 4 below. The Birds of North America account (Mowbray,
et al. 2002) includes a summary and recognizes 11 subspecies: Canada Goose Branta canadensis - Large bodied group - 7 subspecies nesting inland and southerly |
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| Cackling Goose Branta hutchinsii - Small-bodied group - 4 subspecies breeding mainly in tundra | ||
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| In all
subspecies, there are subpopulations and gradual clinal changes so that it
is difficult to define the characteristics of that subspecies without
looking at a broad geographic sample. In addition, the distribution of
Canada Geese is dynamic, changing frequently and rapidly in response to
climate, food resources, and other, often man-made changes. The Cackling
Cackling Goose B. h. minima which formerly wintered almost exclusively in the Central Valley of California,
shifted its winter range northward in the 1990s and now winters mainly
in western Oregon. The history of land use in North America over the last
300 years, and the recent release and introduction of large Canada Geese
across the lower 48 states, has obliterated any geographic variation and
natural distribution patterns that existed before European settlement of
these areas. We can anticipate further rapid change in range and potential
mixing of populations.
Summarizing the subspecies |
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| Bill
size Actual and proportional bill length may be the single most useful feature to study when trying to identify Canada and Cackling Geese (see Figure 1). In general the smaller subspecies have shorter, stubbier, proportionally smaller bills, while the larger subspecies have longer, proportionally larger bills. Unfortunately there seems to be overlap between the smaller populations of Canada Goose and larger subspecies of Cackling Goose. ![]() Figure 1. Mean culmen length of 11 subspecies of Canada and Cackling Geese (from Mowbray et al 2002 and Leafloor et al 1998). The four on the left are included in the new Cackling Goose, while the nine on the right make up the new, restricted, Canada Goose. The two samples on the far right from Akimiski Island are the subspecies interior, and show data from Leafloor et al., demonstrating that captive-raised birds grew larger than similar birds raised in the wild there. Another way of looking at these numbers, and perhaps a better indication
of how useful bill length might be for distinguishing subspecies in the
field, is shown here in figure 2. Size
Compounding the problem of distinguishing subtle size differences, it has been shown that environmental factors (probably nutrition) can dramatically influence the overall size of individual geese. Thus birds raised in environments with reduced quality and/or quantity of food (such as some northern goose nesting colonies that are overcrowded) fail to reach their full potential size, and grow into relatively small geese regardless of their genetic background. Leafloor et al (1998) report that captive-raised birds from Akimiski Island in James Bay have culmen length averaging 5 to 8% larger than wild birds from the same nesting area. While this is certainly a large enough difference to be visible in the field, and should be considered when judging size of geese, it may have been overstated in recent identification discussions. Judging from the data here, the difference between the "runt" wild-raised birds from Akimiski Island and their better-nourished counterparts is not enough to cause confusion of these small Canada Geese B. c. interior with Cackling Geese B. h. hutchinsii, for example (see Table 2). The four to six percent difference in size is about the same as the difference between males and females within each subspecies and is less than the natural variation within each subspecies (one standard deviation equals plus or minus about 4 to 8%, with further variation geographically; data from Mowbray et al 2002). Leafloor et al report that the "runt" birds from Akimiski Island match the size of the same subspecies from 600 km north in Manitoba, tracking the gradual decrease in size northward. Leafloor et al do report some differences in proportions, primarily that while captive-raised adults have culmen length 5 to 8% larger, tarsus length is 11 to 14% larger than wild birds. This is not enough data from which to draw any firm conclusions, at least it shows that the tarsus becomes proportionally smaller than the bill in "runt" wild birds, and suggests that these birds might actually appear proportionally large-billed. There is no evidence that the "runt" birds from Akimiski Island take on the short-billed proportions of smaller subspecies, and therefore should not cause confusion with Cackling Goose. A website shows photos from Michigan of a banded goose from Akimiski Island that is smaller than its associates. One photo shows the bird looking very small, while another shows the expected subtle difference. The accompanying discussion implies that Akimiski Island birds are small and dark, but no color differences are mentioned by Leafloor et al (1998). I don't know if there is evidence of such "environmental darkening". Neck
length Overall
color White
collars Black chin
stripes Voice Wing
shape The
identification problem Distinguishing B. h. hutchinsii from large Canada Geese is relatively easy. In general hutchinsii is much smaller with a short, stubby bill, relatively round or even square head with steep forehead (vs wedge-shaped head), and relatively short neck. They appear paler silvery gray on the upperparts and upperwing coverts in some lighting conditions, with the feathers having a more conspicuous dark subterminal bar and a broader and more conspicuous pale tip. The breast color averages slightly darker than large eastern Canadas, although many are pale-breasted, and the breast often has a golden wash. The voice is slightly higher-pitched, and the wing shape in flight is different. On the other hand, distinguishing B. h. hutchinsii from smaller individuals of Lesser Canada Goose B. c. parvipes may be much more difficult. Many Lesser Canada Geese are noticeably larger than hutchinsii, with proportionally large bills approaching the large Canada Geese, but Lessers from the northern parts of their breeding range are reportedly smaller than in the south. More study is needed to figure out the characteristics of small Lesser compared to hutchinsii. Taverner's is simply poorly known, with a range in western and northern Alaska between the range of Lesser Canada Goose and true Cackling (B. h. minima). It reportedly differs from Lesser in mtDNA, suggesting little interbreeding, but differences in appearance are few and subtle, so that field studies generally claim widespread interbreeding and the entire population on the North Slope of Alaska is listed by Mowbray et al as taverneri/parvipes. Clearly, this is another topic for further study. Finally, as a cautionary note, Pearce et al (2000) used genetic testing to assess the accuracy of identifications of Dusky Canada Geese B. c. occidentalis at hunter check stations in Washington. They found that one-third to one-half of all the geese identified as Dusky by breast color and bill length (measured in the hand) were in fact not true Duskys. And if that's the level of accuracy achieved by experienced people with birds in hand, what can we expect to do in the field? This is an irresistible challenge for some of us, but until we have a better sense of the variation and limits of these subspecies all identification should be approached with a healthy dose of skepticism.
Leafloor, J. O., C. D. Ankney, and D. H. Rusch. 1998. Environmental effects on body size of Canada Geese. Auk 115:26-33, Marquardt, R. E, 1961. Albinism in the small white-cheeked geese. Auk 78:99-100 Mowbray, T. B., C. R. Ely, J. S. Sedinger, and R. E. Trost. 2002. Canada Goose (Branta canadensis). In The Birds of North America, No. 682 (A. Poole and F. Gill, eds.). The Birds of North America, Inc., Philadelphia, PA. Palmer, R. S., ed. 1976. Handbook of North American Birds; volume 2, Waterfowl (Part 1). Yale University Press. Pearce, J. M., B. J. Pierson, S. L.Talbot, D. V. Derksen, D. K. Kraege, and K. T. Scribner. 2000. A Genetic Evaluation of Morphology Used to Identify Harvested Canada Geese. Journal of Wildlife Management 64(3): 863-874. Sibley, D. A. 2000. National Audubon Society The Sibley Guide to Birds. Chanticleer Press, NY. Whitford, P. C. 1998. Vocal and visual communication of Giant Canada Geese. Pp. 375-386 in Biology and management of Canada Geese (D. H. Rusch, M. D. Samuel, D. D. Humburg, and B. D. Sullivan, eds.). Proc. Intl. Canada Goose Symp., Milwaukee, WI. Other references:
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Figure 4. Map showing basic range of subspecies of Canada and Cackling Goose. Cackling Goose breeding range in darker pink-red; Canada Goose breeding range in paler orange-red. Winter range of all subspecies in blue, gray lines connect some breeding populations to their principal winter range. Many populations engage in a mid-summer molt migration, traveling up to hundreds of miles north of their breeding range. (based on Mowbray et al 2002 and some comments from Ian Mclaren). |
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