| Olive-sided Flycatcher Contopus borealis
Northern
Pacific
I've distinguished these two populations based on differences in song between coastal northern California and Taiga breeders, but this geographic variation does not quite match with the currently named subspecies. Southern California breeders (C. b. marjorinus) average slightly larger with darker underparts than the widespread subspecies (C. b. borealis/cooperi??). Obviously more study is needed to reconcile the named subspecies with variation in song.
Willow Flycatcher Empidonax traillii
Eastern E. t. traillii group
Southwestern E. t. brewsteri group
Variation in this species is complex and extremely subtle, with many published revisions by different researchers over the last 50 years. No differences in voice are known, the differences in plumage are clinal and variable.
Pacific Slope Flycatcher Empidonax difficilis
Mainland E. d. difficilis
Channel Islands E. d. insulicola
Brown-crested Flycatcher Myiarchus tyrannulus
Arizona M. t. magister
Texas M. t. cooperi
Tropical Kingbird Tyrannus melancholicus
Western T. m. occidentalis
Eastern T. m. satrapa
These two populations differ very slightly in appearance according to Pyle: Eastern birds have a slightly deeper tail fork on average, and slightly richer colors overall. Possible differences in voice that are described in the guide need more study.
Fork-tailed Flycatcher Tyrannus savana
South American T. s. savana group
Central American T. s. monachus
Eurasian Skylark Alauda arvensis
European A. a. arvensis
Asian A. a. pekinensis
The introduced population in British Columbia originated in the British Isles (A. a. arvensis), but it is quite possible that plumage features have changed during the subsequent xx years in the very humid environment of Vancouver Island. Vagrants in Alaska (and once in California) are the Asian subspecies A. a. pekinensis.
Horned Lark Eremophila alpestris
West Arctic E. a. arcticola
Pacific Northwest E. a. strigata
Interior West E. a. occidentalis
Central California E. a. sierrae
Southeast Arizona E. a. adusta
South Texas E. a. giraudi
Northeast E. a. alpestris
These subspecies are examples of the range of variation found across North America, where 21 subspecies are listed in Pyle 1997. There are trends in the variation as described in the book, but not clear groups of subspecies.
Purple Martin Progne subis
Eastern P. s. subis
Western P. s. hesperia group
These two populations differ in voice, size, plumage, migration timing, and do not overlap in range.
Cliff Swallow Hirundo pyrrhonota
Northern H. p. pyrrhonota
Southwestern H. p. swainsoni
These two species differ in size and plumage color. Two other subspecies in the southwest are intermediate to some extent, and all subspecies blend at edges of their range.
Cave Swallow Hirundo fulva
Mexican H. f. pelodoma
Caribbean H. f. fulva
These two populations do not overlap in breeding range. They differ in size and slightly in overall plumage color. Possible differences in voice need more study.
Barn Swallow Hirundo rustica
American H. r. erythrophthalmus group
Eurasian H. r. rustica group
Eurasian subspecies rustica and gutturalis have both been identified in Alaska. American population is difficult to distinguish, as the underparts color varies from whitish to rusty.
Gray Jay Perisoreus canadensis
Taiga P. c. canadensis
Rocky Mts P. c. capitalis
Pacific P. c. obscurus
Several other subspecies are intermediate between these three populations.
Steller's Jay Cyanocitta stelleri
Pacific C. s. stelleri group
Queen Charlotte Islands C. s. carlottae
Interior C. s. macrolopha group
Blue Jay Cyanocitta cristata
Northern C. c. bromia
Southern C. c. cristata
These two subspecies more or less match the characteristics illustrated in the book, but their distribution does not match my limited observation. The population at Cape May, NJ seems to be subtly dimorphic, including both pale and dark blue birds, while birds in the Adirondacks are darker blue than even the darkest NJ birds. More study is needed.
Western Scrub Jay Aphelocoma californica
Pacific A. c. californica group
Interior West A. c. woodhouseii group
Gray-breasted Jay Aphelocoma ultramarina
Texas A. u. couchii
Arizona A. u. arizonae
Black-billed Magpie Pica hudsonia
No subspecies are described within North America, although subtle clinal size differences are known. The Northern and Southern examples illustrated may simply represent the ends of clinal variation. More study is needed.
American Crow Corvus brachyrhynchos
Currently accepted subspecies are based on subtle clinal variations in size and do not match the vocal differences described in the book. Populations from Washington to California (and perhaps farther east as well) sound distinctly different from Atlantic coast birds.
Black-capped Chickadee Parus atricapillus
Eastern P. a. atricapillus group
Rocky Mountain P. a. septentrionalis group
Pacific P. a. occidentalis group
Carolina Chickadee Parus carolinensis
Northeast P. c. extimus
Southwest P. c. agilis
These two subspecies represent the extremes of color variation in this species. Other subspecies are intermediate.
Mountain Chickadee Parus gambeli
Pacific P. g. baileyae
Rocky Mountains P. g. gambeli
Differences between these subspecies are slight, and there are several other intermediate subspecies in the interior west.
Boreal Chickadee Parus hudsonicus
Alaska P. h. columbianus group
Typical P. h. hudsonicus group
Chestnut-backed Chickadee Parus rufescens
Typical P. r. rufescens
Central California Coast P. r. barlowi
Marin County, CA is occupied by an intermediate subspecies P. r. neglectus.
Tufted Titmouse Parus bicolor
Tufted P. b. bicolor
Black-crested P. b. atricristatus group
Bushtit Psaltriparus minimus
Pacific P. m. minimus group
Interior P. m. plumbeus group
Black-eared P. m. melanotis group (the subspecies occurring in North America is P. m. dimorphicus).
White-breasted Nuthatch Sitta carolinensis
Eastern S. c. carolinensis group
Mountain S. c. nelsoni group
Pacific S. c. aculeata group
Pygmy Nuthatch Sitta pygmaea
Interior S. p. melanotis
Central California S. p. pygmaea
Brown Creeper Certhia americana
Eastern C. a. americana group
Western C. a. occidentalis group
Mexican C. a. albescens
Cactus Wren Campylorhynchus brunneicapillus
Interior C. b. couesi group
Coastal Southern California C. b. affine group (the subspecies found in California is C. c. sandiegense).
Carolina Wren Thyrothorus ludovicianus
Mexican T. l. lomitensis
Northern T. l. ludovicianus group
Bewick's Wren Thryomanes bewickii
Eastern T. b. bewickii
Southwestern T. b. eremophilus group
Pacific T. b. drymoecus group
House Wren Troglodytes aedon
Eastern T. a. aedon
Western T. a. parkmanii
Brown-throated T. a. cahooni group (the intermediate birds in Arizona have been given the name T. a. 'vorhiesi')
Winter Wren Troglodytes troglodytes
Eastern T. t. hiemalis group
Pacific T. t. pacificus group
Pribilofs T. t. alascensis group
Marsh Wren Cistothorus palustris
Eastern C. p. palustris group
Worthington’s C. p. griseus
Western C. p. paludicola group
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