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Northern Fulmar
Fulmarus glacialis
Atlantic F. g. glacialis group
Pacific F. g. rodgersii
Atlantic and Pacific populations do not overlap in distribution and
are apparently separable by plumage, they also differ slightly in size
and bill size, although these differences are complicated by variation.
Within each ocean there are small variations in size, proportions, and
color (frequency of color morphs). Within this mosaic of subtle variation
anywhere from one to three subspecies are recognized in the North Atlantic.
Cory's Shearwater
Calonectris diomedea
Mediterranean C. d. diomedea
Atlantic C. d. borealis
Two subspecies differ in size and subtly in plumage but are very difficult
to distinguish, perhaps indistinguishable at sea. Mediterranean birds
have been recorded off New York. Atlantic island breeders apparently
account for the vast majority of all North American records. These two
populations have been proposed as full species. The more distinctive
Cape Verde Islands population C. d. edwardsii (often considered
a separate species Cape Verde Shearwater C. edwardsii) is unconfirmed
in North America.
Leach's Storm-Petrel
Oceanodroma leucorhoa
White-rumped O. l. leucorhoa
Dark-rumped O. l. chapmani
Variation is complex, the named subspecies are not always white- or
dark-rumped as implied here. Taxonomy was revised by Ainley (1980. Auk
97:837-853). Nominate O. l. leucorhoa is widespread in the north,
usually white-rumped, and individuals from the North Atlantic are indistinguishable
from those in the North Pacific. The subspecies O. l. chapmani differs
from leucorhoa in averaging smaller in size, in some very subtle
differences in proportions, and in being mainly dark-rumped (few leucorhoa
are dark-rumped), but do not differ significantly in voice. They
apparently differ in foraging habitat preference, and may differ in
other ways as well, but all differences are clinal with many intermediate
birds off central and southern California. Two other populations nest
on Guadalupe Island off Mexico and could wander to waters off California.
The winter breeding population there is named O. l. cheimomnestes
and the summer breeding population O. l. socorroensis.
Brown Booby
Sula leucogaster
Atlantic S. l. leucogaster
Eastern Pacific S. l. brewsteri
These two subspecies apparently differ only in adult male plumage, specifically
in head color.
Red-footed Booby
Sula sula
Caribbean S. s. sula
Eastern Pacific S. s. websteri
Two subspecies differ in adult plumage, with most or all adult S.
s. sula white-tailed and most or all adult websteri dark-tailed.
No other differences reported, and other subspecies in Pacific and Indian
oceans also vary in tail color.
Brown Pelican
Pelecanus occidentalis
Atlantic P. o. occidentalis group
Pacific P. o. californicus group
Two subspecies differ slightly in size and breeding colors. Many are
identifiable in the field. Interestingly, band recoveries indicate quite
a bit of movement between these populations across Central America.
Double-crested
Cormorant Phalacrocorax auritus
Eastern P. a. auritus group
Western P. a. cincinatus group
Subspecies differ clinally in size and in extent of white in breeding
plumes but no other significant differences, any division is arbitrary.
Pelagic Cormorant
Phalacrocorax pelagicus
Southern P. p. resplendens
Bering Sea P. p. pelagicus
Northern birds average larger than Southern. The arbitrary division
between these subspecies is around the Queen Charlotte Islands, BC,
but variation is clinal with much overlap. The population in the Bering
Sea may be more distinctive.
Great Blue Heron
Ardea herodias
Great Blue A. h. herodias group
Great White A. h. occidentalis
Variation in this species is particularly interesting. Great White Heron
was formerly considered a separate species distinguished from Great
Blue by plumage color, size, and length of head plumes. Great White
is known to interbreed with dark (Great Blue) birds within its range,
and the intermediate "Wurdemann's" Heron (initially described as a separate
species) was considered a hybrid. Now all of these southern Florida
birds are considered white, intermediate, and dark morphs of a distinctive
subspecies A. h. occidentalis, characterized by large size and
short head plumes.
Greater Flamingo
Phoenicopterus ruber
American P. r. roseus
Eurasian P. r. ruber
Careful readers will notice that this is the only Latin subspecies name
included in the book, simply because the way the page is laid out it
seemed best to include the name for clarity. In fact I believe these
are two distinct species with differences in plumage, bare parts colors,
voice, and with no range overlap.
Tundra Swan
Cygnus columbianus
Whistling C. c. columbianus
Bewick's C. c. bewickii
Two subspecies distinguished by extent of yellow on bill. Sometimes
considered separate species, but there is near overlap in bill pattern
and no other differences known.
Greater White-fronted
Goose Anser albifrons
Tundra A. a. frontalis
Taiga A. a. gambelli
Tule A. a. elgasi
Greenland A. a. flavirostris
Variation is subtle and complex, although both Greenland and Tule populations
have restricted range and are fairly distinctive in size, plumage, and
bill color. The variation within Tundra and Taiga populations creates
extensive overlap.
Snow Goose
Chen caerulescens
Lesser C. c. caerulescens
Greater C. c. atlanticus
Two subspecies differ in size and in frequency of dark morph. Most can
be identified but there is considerable overlap.
Brant
Branta bernicla
Black B. b. nigricans
Intermediate B. b. unnamed?
Pale-bellied B. b. hrota
Three subspecies in North America differ subtly in plumage. The status
and identification of the recently diagnosed Intermediate population,
nesting on Melville Island and wintering in Puget Sound, is poorly known.
[Eurasian race B. b. bernicla could occur in North America, but
would probably be indistinguishable from Intermediate birds and from
Black X Pale-bellied intergrades].
Canada Goose
Branta canadensis
Common B. c. canadensis group
Lesser B. c. parvipes
Richardson's B. c. hutchinsii
Dusky B. c. occidentalis group
Cackling B. c. minima
Aleutian B. c. leucopareia
Rather arbitrarily divided here into six subspecies groups that differ
in size, plumage, and voice; but all are connected by intermediate populations
and identification can be difficult, even though the extremes seem like
different species.
Green-winged
Teal Anas crecca
American A. c. carolinensis
Eurasian A. c. crecca group
Two populations sometimes considered separate species. They differ only
in adult male plumage. Interbreeding occurs in the limited area where
range overlaps.
Mallard
Anas platyrhynchos
Northern A. p. platyrhynchos group
Mexican A. p. diazi
Two subspecies differ in plumage, especially adult males. Identification
and status in North America is complicated by the fact that virtually
no pure "Mexican" Mallards occur; all show evidence of intergradation
with Northern Mallards.
Common Eider
Somateria mollissima
West Arctic S. m. v-nigra
Hudson Bay S. m. sedentaria
East Arctic S. m. borealis
Atlantic S. m. dresseri
Four subspecies differ in bill structure, adult male bill color, and
subtly in female color. Variation within each subspecies makes identification
difficult, and most populations intergrade where ranges meet (and it
is possible that other subspecies could wander to our northeastern shores
from Iceland or Europe). West Arctic and Hudson Bay populations are
more or less isolated. West Arctic population is the most distinctive
in appearance and has been proposed for species status. More research
is needed on field identification.
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