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Western Spindalis Spindalis zena
Green-backed S. z. ??
Black-backed
Summer Tanager Piranga rubra
Eastern P. r. rubra
Western P. r. cooperi group
Northern Cardinal Cardinalis cardinalis
Southwestern C. c. superbus
Eastern C. c. cardinalis group
Painted Bunting Passerina ciris
Eastern P. c.
Western P. c.
Eastern Towhee Pipilo erythrophthalmus
Red-eyed P. e. erythrophthalmus group
White-eyed P. e. alleni
Spotted Towhee Pipilo maculatus
Pacific Northwest P. m. oregonus group
Southwest P. m. montanus group
Great Plains P. m. arcticus
Bachman's Sparrow Aimophila aestivalis
Western A. a. illinoensis
Eastern A. a. aestivalis
Botteri's Sparrow Aimophila botterii
Texas A. b. texana
Arizona A. b. arizonae
Rufous-crowned Sparrow Aimophila ruficeps
Pacific A. r. ruficeps group
Interior A. r. scottii group
American Tree Sparrow Spizella arborea
Pale S. a. ochracea
Dark S. a. arborea
Brewer's Sparrow Spizella breweri
Southern S. b. breweri
Northern (Timberline) S. b. taverneri
Field Sparrow Spizella pusilla
Gray S. p. arenacea
Rufous S. p. pusilla
Black-throated Sparrow Amphispiza bilineata
Texas A. b. bilineata
Intermediate A. b. opuntia
Western A. b. deserticola
Sage Sparrow Amphispiza belli
Interior A. b. nevadensis
Pacific (Bell's) A. b. belli
These two subspecies are easily distinguished by plumage and song, and species status has been proposed. The situation is complicated somewhat by an intermediate subspecies, A. b. canescens, in interior California.
Savannah Sparrow Passerculus sandwichensis
Typical P. s. sandwichensis group
Ipswich P. s. princeps
Belding's P. s. beldingi
Aleutian P. s. sandwichensis group
Large-billed Sparrow P. s. rostratus group
Variation in this species is considerable with a number of named subspecies across North America. Most variation is clinal, however, and only a few subspecies and subspecies groups are really distinguishable. In general subspecies in the east and in humid Pacific coastal areas are more rufous, while those in the drier interior are gray, but there is considerable variation and I found it difficult to separate broad eastern and western groups among the continental birds. Of these the most distinctive in plumage, structure, voice and DNA is the Large-billed group, which has been proposed for species status. In the past Ipswich has been considered a separate species, a treatment supported by its isolation during the breeding season and clearly different size and plumage.
Grasshopper Sparrow Ammodramus savannarum
Northern A. s. pratensis group
Florida A. s. floridanus
Within the Northern pratensis group there are several subspecies, the most distinctive of which is the relatively pale and rufous Arizona population A. s. ammolegus.
Nelson’s Sharp-tailed Sparrow
Interior A. n. nelsoni/alterus
Atlantic A. n. subvirgatus
Seaside Sparrow Ammodramus maritimus
Atlantic A. m. maritimus group
Gulf Coast A. m. sennetti group
Cape Sable A. m. mirabilis
[Dusky] A. m. nigrescens
Fox Sparrow Passerella iliaca
Red (Taiga) P. i. iliaca group
Sooty (Pacific) P. i. unalaschensis group
Slate-colored (Interior West) P. i. schistacea group
Thick-billed (California) P. i. megarhyncha group
Song Sparrow Melospiza melodia
Eastern M. m. melodia
Pacific Northwest M. m. morphna
Aleutian M. m. maxima
Central California coast M. m. gouldii
Southwestern M. m. saltonis
There are 29 named subspecies of Song Sparrow listed in Pyle, and I have tried to show the range of variation by illustrating these five subspecies, but all variation is clinal and it is difficult to categorize the other 24 subspecies into groups.
White-crowned Sparrow Zonotrichia leucophrys
Eastern Taiga Z. l. leucophrys
Western Taiga Z. l. gambellii
Interior West Z. l. oriantha
Pacific Z. l. nuttalli group (includes Z. l. pugetensis)
Dark-eyed Junco Junco hyemalis
Slate-colored J. h. hyemalis group
Canadian Rocky Mountains J. h. cismontanus
Oregon J. h. montanus group
Pink-sided J. h. mearnsi
Gray-headed J. h. caniceps
Red-backed J. h. dorsalis
White-winged J. h. aikeni
Red-winged Blackbird Agelaius phoeniceus
Typical A. p. phoeniceus group
Bicolored A. p. californicus group
This is another complex species, with 14 named subspecies. Variation is somewhat clinal and there are several named subspecies intermediate between these two. Identification may be difficult or impossible in practice.
Eastern Meadowlark Sturnella magna
Typical S. m.
Lilian’s (Southwestern) S. m. lilianae
Great-tailed Grackle Quiscalus mexicanus
Eastern Q. m. prosopidicola
Western Q. m. nelsoni
Boat-tailed Grackle Quiscalus major
Atlantic Q. m. torreyi
Gulf Coast/ Florida Q. m. major group
Common Grackle Quiscalus quiscula
Interior (Bronzed) Q. q. versicolor
Coastal (Purple) Q. q. stonei
Florida Q. q. quiscula
Bronzed Cowbird Molothrus aeneus
Eastern M. a. aeneus
Western M. a. loyei
Brown-headed Cowbird Molothrus ater
Eastern M. a. ater
Pacific/Mexican M. a. obscurus
The intermediate population is named M. a. artemisiae.
Hooded Oriole Icterus cucullatus
Western I. c. nelsoni
Eastern I. c. sennetti
The intermediate population is named I. c. cucculatus.
Gray-crowned Rosy Finch Leucosticte arctoa
Interior (Gray-crowned) L. a. tephrocotis group
Coastal (Gray-cheeked or Hepburn's) L. a. littoralis group
Bering Sea L. a. griseonucha group
Pine Grosbeak Pinicola enucleator
Pacific P. e. flammula group
Interior West P. e. montanus group
Taiga P. e. leucurus
Purple Finch Carpodacus purpureus
Eastern C. p. purpureus
Pacific C. p. californicus
Red Crossbill Loxia curvirostra
The current state of crossbill taxonomy in the northern hemisphere defies subspecies names. Nine different call types corresponding to subtle differences in bill size have been diagnosed in North America (and recent research in Europe has uncovered a similar situation there). These populations have been designated by numbers or letters - e.g. types 1-9 or a-i - and for the most part do not correspond to previously named subspecies. Under the circumstances I think it is simplest to ignore all previous subspecies names and refer to these birds simply by the number or letter designations, rather than trying to reconcile the two systems of naming. More information can be found in the technical literature.
Common Redpoll Carduelis flammea
Southern C. f. flammea
Greenland (Greater) C. f. rostrata
Hoary Redpoll Carduelis hornemanni
Southern C. h. exilipes
Greenland C. h. hornemanni
Lesser Goldfinch Carduelis psaltria
Western C. p. psaltria
Texas C. p. mexicanus
Evening Grosbeak Coccothraustes vespertinus
Eastern C. v. vespertinus
Western C. v. montanus group
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